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The body can be the only difference.
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Source of the two once fired cases?
.308 Winchester -Two once fired cases with IDENTICAL head space measurement and OAL after FL sizing and trimming to 2.005". One chambers, the other doesn't.
Cases are Federal GMM brass, two different lots.
Any ideas???
And, before using a small base die, please check for impending "Incipient Case Head Separation";
https://www.youtube.com/watch?v=dppyuA0lmYU
UPDATE!!
So I took BearNaked's advice and got my Sharpie out. After coloring all of the case but the neck and chambering the case there's a bright ring that circumscribes the case at the web.
I also noticed that when just inserting the same case into the chamber it sticks with just finger pressure without even closing the bolt. So what's the concensus? Body die? Small base body die? Or try getting more of the case up into a regular FL die?