To simplify this question and answer I will refer to rimfire actions/barrels only.... Until the RimX, gunsmiths had to have the action in hand to properly headspace the barrel.... This is very very important for rimfire accuracy as the headspace typically matched the ammo the customer was shooting (rim thickness). Its different for Lapua vs say Eley, etc... If you had excessive headspace like say .049" and were shooting Eley Tenex with a 0.039" rim thickness, your bullet was not consistently tight against the breach face on every round so you lost your consistency. This is because the extractors only held the case till the bolt was closed at which time the extractors spread apart via the extractor grooves so the bolt face is the only thing holding that bullet in the chamber.. If too much headspace, you have bullet movement...
With this RimX, Zermatt has a tenon print they provide to smiths so gunsmiths can cut barrels to Zermatt's tenon print and because Zermatt's CNC machining on these actions are so tight and so consistent, every action is the same. So when a smith makes barrel after barrel to Zermatt's tenon print spec they fit every RimX... The key here is consistency from action to action which is what makes this possible.
Now the headspace thing....Zermatt uses a tensioner that once the bolt is slid up onto the bolt face all the way, the tensioner grabs and holds the case rim against the bolt face.... So no matter what, the case rim is consistently in the same place, shot after shot...The case cant move with creates consistency. Consitency from shot to shot is key with 22LR. So now your only inconsistency is ammo which has always been the weakest link with 22LR... This is why we buy quality ammo and lot test, sort, etc...
The RimX does come with this little headspace gauge you can use to check to make sure youre good. Honestly, havent used it to date but one shop is doing my barrels and they are 100% exactly the same. The RimX has a minimum headspace of 0.0455" and maximum of 0.050". I have Keystone setting headspace to 0.046" on every one of my barrels..
@Zermatt Arms can jump in here if I missed anything or was off on anything but I believe I explained it best as possible.